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Batuwita S, Udugampala S, De Silva M, Diao J, Edirisinghe U. A review of amphibian fauna of Sri Lanka: distribution, recent taxonomic changes and conservation. JAD 2019; 1 (2) :44-82
URL: http://jad.lu.ac.ir/article-1-45-en.html
1- The Society for the Biodiversity Conservation, 63/4, Adikaramwatta, Yaggahapitiya, Gunnepana 20270, Kandy, Sri Lanka , sudesh.batuwita@gmail.com
2- Wildlife Conservation Society- Galle, Hiyare reservoir, Hiyare 80000, Galle, Sri Lanka
3- College of Biology and Environment, Nanjing Forestry University, 159 Longpan Road, Nanjing, China
4- Postgraduate Institute of Agriculture, University of Peradeniya, Peradeniya 20200, Sri Lanka
Abstract:   (24790 Views)
The amphibian fauna of Sri Lanka comprises 120 species, including 107 (~90.0%) endemic species. They belong to five families: Bufonidae, Dicroglossidae, Ichthyophiidae, Microhylidae, and Rhacophoridae. Based on distribution, we recognized five zoogeographic zones for them, Central Hills, Dry Zone, Knuckles Range, Lowland Wet Zone, and Rakwana Hills. Fifty three species were reported from the Central Hills (48 endemics [90.6%] and 42 [79.2%] threatened species). 47 species were recorded from the Lowland Wet Zone, including 36 (76.6%) endemics and 28 (59.6%) threatened species. The Knuckles Range had 25 species, of which, 19 (76.0%) were endemics and 15 (60.0%) are threatened species. 19 species were reported from Dry Zone including seven endemics (36.8%) and four threatened species (21.1%). Out of 29 species, which inhabited in the Rakwana Hills, 26 were endemics (~89.7%) including 24 (82.8%) threatened species. Species diversity along the elevational gradient was also observed with the highest species richness in the mid-elevational localities. Family Ichthyophiidae can be considered as the least studied family. Recent rediscoveries and studies have helped to reduce the number of extinct species from 21 to 18. It is speculated that some of the other extinct species have to be rediscovered or probably were misidentified as other species. About 90% of Sri Lankan amphibians occur in the regions with the highest human populations where there are established agricultural lands. Loss of habitats, competition due to anthropogenic species and invasive species, pollution (cause for malformations, parasites, and other diseases), and climate change appear to be major threats.
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Type of Study: Original Research Article |
Received: 2019/11/25 | Accepted: 2019/12/24 | Published: 2019/12/31

References
1. Ahl, E. (1927). Zur systematik der asiatischen arten der frosch gattung Rhacophorus. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1927: 35–47.
2. Alam, M. S., Igawa, T., Khan, M. M. R., Islam, M. M., Kuramoto, M., Matsui, M., Kurabayashi, A. and Sumida, M. (2008). Genetic divergence and evolutionary relationships in six species of genera Hoplobatrachus and Euphlyctis (Amphibia: Anura) from Bangladesh and other Asian countries revealed by mitochondrial gene sequences. Molecular Phylogenetics and Evolution, 48 (2): 515–527. https://doi.org/10.1016/j.ympev.2008.04.020 [DOI]
3. Amarasekara, M. G. T. S., Dayawansa, N. D. K. and De Silva, R. P. (2013). Stream water quality depletion under intensive agriculture. South Asian Water Studies, 4 (1): 23–37.
4. Anderson, J. (1871). A list of the reptilian accession to the Indian Museum, Calcutta from 1865 to 1870, with a description of some new species. The Journal of the Asiatic Society of Bengal, 40: 12–39.
5. Annandale, N. (1913). Some new and interesting Batrachia and lizards from India, Ceylon and Borneo. Records of the Indian Museum, 9: 301–307.
6. Annandale, N. (1919). The fauna of certain small streams in the Bombay Presidency: some frogs from streams in the Bombay Presidency. Records of the Indian Museum, 16: 109–161.
7. Anonymous (1995). Sri Lanka Forestry Sector Master Plan. Forestry planning unit, Ministry of agriculture, land and forestry, Colombo, Sri Lanka. 511 pp.
8. Anonymous (2003). Statistical Pocket Book 2003. Department of Census and Statistics, Colombo, Sri Lanka. 100 pp.
9. Ashton, M. S., Gunatilleke, S., de Zoysa, N., Dassanayake, M. D., Gunatilleke, N. and Wijesundera, S. (1997). A field guide to the common trees and shrubs of Sri Lanka. WHT Publications (Pvt.) Ltd, Colombo, Sri Lanka. 432 pp.
10. Bahir, M. M. and Silva, A. (2005). Otocryptis nigristigma, a new species of agamid lizard from Sri Lanka. The Raffles Bulletin of Zoology, 12: 393–406.
11. Bahir, M. M. and Surasinghe, T. (2005). A conservation assessment of the agamid lizards of Sri Lanka. The Raffles Bulletin of Zoology, 12: 407–412.
12. Bahir, M. M., Meegaskumbura, M., Manamendra-Arachchi, K., Schneider, C. J. and Pethiyagoda, R. (2005). Reproduction and terrestrial direct development in Sri Lankan shrub frogs (Ranidae: Rhacophorinae: Philautus). The Raffles Bulletin of Zoology, 12: 339–350.
13. Bambaradeniya, C. N. B. (1999). Alien invasive fauna in natural habitats and their impact on indigenous biota, In: Marambe, B. (Ed.), Proceedings of the First National Workshop on Alien Invasive Species in Sri Lanka. Ministry of Forestry and Environment, Sri Lanka. pp. 45–51.
14. Basnayake, B. R. S. B. (2007). Climate change, In: Anonymous, The National Atlas of Sri Lanka. Government Press Sri Lanka, Colombo. pp. 54–55.
15. Batuwita, S. (2000). Lizards of Mount Gannoruwa. Loris, 22 (3): 13–15.
16. Batuwita, S. (2019). A review of the lizards of the endemic genus Lankascincus (Reptilia: Scincidae: Lygosominae) from Sri Lanka. Bulletin of the Museum of Comparative Zoology, 162 (3): 211–262. https://doi.org/10.3099/MCZ38.1 [DOI]
17. Batuwita, S. and Bahir, M. M. (2005). Description of five new species of Cyrtodactylus (Reptilia: Gekkonidae) from Sri Lanka. The Raffles Bulletin of Zoology, 12: 351–380.
18. Batuwita, S. and Udugampala, S. (2017). Description of a new species of Cnemaspis (Squamata: Gekkonidae) from Knuckles Range of Sri Lanka. Zootaxa, 4254 (1): 82–90. http://doi.org/10.11646/zootaxa.4254.1.4 [DOI]
19. Batuwita, S., Agarwal, I. and Bauer, A. M. (2019a). Description of a new diminutive, rupicolous species of day-gecko (Squamata: Gekkonidae: Cnemaspis) from southern Sri Lanka. Zootaxa, 4565 (2): 223–234. http://dx.doi.org/10.11646/zootaxa.4565.2.6 [DOI]
20. Batuwita, S., De Silva, M. and Udugampala, S. (2019b). Description of a new species of Pseudophilautus (Amphibia: Rhacophoridae) from southern Sri Lanka. Journal of Threatened Taxa, 11 (1): 13120–13131. https://doi.org/10.11609/jott.3903.11.1.13120-13131 [DOI]
21. Bierregaard, R. O., Gascon, C., Lovejoy, T. E. and Masquita, R. (2001). Lessons from Amazonia: the ecology and conservation of a fragmented forest. Yale University Press, New Haven, Connecticut, and London, United Kingdom. 496 pp.
22. Biju, S. D., Garg, S., Mahony, S., Wijayathilaka, N., Senevirathne, G. and Meegaskumbura, M. (2014). DNA barcoding, phylogeny and systematics of Golden-backed frogs (Hylarana, Ranidae) of the Western Ghats-Sri Lanka biodiversity hotspot, with the description of seven new species. Contributions to Zoology, 83 (4): 269–335. https://doi.org/10.1163/18759866-08304004 [DOI]
23. Biju, S. D., Shouche, Y. S., Dubois, A., Dutta, S. K. and Bossuyt, F. (2010). A ground-dwelling rhacophorid frog from the highest mountain peak of the Western Ghats of India. Current Science, 98: 1119–1125.
24. Blyth, E. (1852). Report of curator, zoological department. The Journal of the Asiatic Society of Bengal, 21: 341–358.
25. Bogart, J. P. (1981). How many times has terrestrial breeding evolved in anuran amphibians? Monitore Zoologico Italiano-Italian Journal of Zoology, 15 (3): 29–40. https://doi.org/10.1080/03749444.1981.10736627 [DOI]
26. Bogert, C. M. and Senanayake, R. (1966). A new species of toad (Bufo) indigenous to southern Ceylon. American Museum Novitates, 2269: 1–18. https://doi.org/10.1080/03749444.1981.10736627 [DOI]
27. Bolkay, S. J. (1915). Beiträge zur Osteologie einiger exotischer Raniden. Anatomischer Anzeiger, Jena, 48: 172–183. https://doi.org/10.5962/bhl.part.17101 [DOI]
28. Bopage, M. M., Wewalwala, K., Krvavac, M., Jovanovic, O., Safarek, G. and Pushpamal, V. (2011). Species diversity and threat status of amphibians in the Kanneliya Forest, lowland Sri Lanka. Salamandra, 47 (3): 173–177.
29. Bossuyt, F. and Dubois, A. (2001). A review of the frog genus Philautus Gistel, 1848 (Amphibia, Anura, Ranidae, Rhacophorinae). Zeylanica, 6 (1): 1–112.
30. Bossuyt, F., Meegaskumbura, M., Beenaerts, N., Gower, D. J., Pethiyagoda, R., Roelants, K., Mannaert, A., Wilkinson, M., Bahir, M. M., Manamendra-Arachchi, K., Ng, P. K. L., Schneider, C. J., Oommen, O. V. and Milinkovitch, M. C. (2004). Local endemism within the Western Ghats-Sri Lanka biodiversity hotspot. Science, 306 (5695): 479–481. https://doi.org/10.1126/science.1100167 [DOI]
31. Boulenger, G. A. (1882a). Catalogue of the Batrachia Salientia s. Ecaudata in the collection of the British Museum. Second Edition. Taylor and Francis, London. xvi+503 pp. https://doi.org/10.5962/bhl.title.12075 [DOI]
32. Boulenger, G. A. (1882b). Description of a new genus and species of frogs of the family Ranidae. Annals and Magazine of Natural History, Series 5, 10 (55): 35. https://doi.org/10.1080/00222938209459662 [DOI]
33. Boulenger, G. A. (1904). Descriptions of three new frogs from southern India and Ceylon. Journal of the Bombay Natural History Society, 15: 430–431.
34. Boulenger, G. A. (1905) [1904]. Description of a frog from Ceylon, hitherto confounded with Rana limnocharis. Spolia Zeylanica, 2: 73–74.
35. Brook, B. W., Sodhi, N. S. and Ng, P. K. L. (2003). Catastrophic extinctions follow deforestation in Singapore. Nature, 424: 420–423. https://doi.org/10.1038/nature01795 [DOI]
36. Caorsi, V. Z., Both, C., Cechin, S., Antunes, R. and Borges-Martins, M. (2017). Effects of traffic noise on the calling behavior of two Neotropical hylid frogs. PLOS ONE, 12 (8): e0183342. https://doi.org/10.1371/journal.pone.0183342 [DOI]
37. Chandramouli, S. R., Ankaiah, D., Arul, V., Dutta, S. K. and Ganesh, S. R. (2019). On the taxonomic status of Minervarya granosa (Kuramoto, Joshy, Kurabayashi and Sumida, 2008) and the distribution of M. agricola (Jerdon, 1853) Amphibia: Anura: Dicroglossidae. Asian Journal of Conservation Biology, 8 (1): 84–87.
38. Clarke, B. T. (1983). A morphological re-examination of the frog genus Nannophrys (Anura: Ranidae) with comments on its biology, distribution and relationships. Zoological Journal of the Linnean Society, 79 (4): 377–398. https://doi.org/10.1111/j.1096-3642.1983.tb01171.x [DOI]
39. Cope, E. D. (1861) [1860]. Descriptions of reptiles from tropical America and Asia. Proceedings of the Academy of Natural Sciences of Philadelphia, 12: 368–374.
40. Dahanukar, N., Sulakhe, S. and Padhye, A. (2017). Identity of Sphaerotheca pluvialis (Jerdon, 1853) and other available names among the burrowing frogs (Anura: Dicroglossidae) of South Asia. Journal of Threatened Taxa, 9 (6): 10269–10285. http://doi.org/10.11609/jott.3358.9.6.10269-10285 [DOI]
41. De Benedictis, P. A. (1974). Interspecific competition between tadpoles of Rana pipiens and Rana sylvatica: an experimental field study. Ecological Monographs, 44 (2): 129–151. https://doi.org/10.2307/1942308 [DOI]
42. De Costa, W. A. J. M. (2008). Climate change in Sri Lanka: myth or reality? Evidence from long-term meteorological data. Journal of the National Science Foundation of Sri Lanka, 36: 63–88. https://jnsfsl.sljol.info/articles/abstract/10.4038/jnsfsr.v36i0.8048/ [DOI]
43. De Silva, A. (1995). The Amphibia of Sri Lanka: a provisional checklist and their common names. Lyriocephalus, 1: 20–26.
44. De Silva, A. (1996). The Amphibia of Sri Lanka: a checklist and an annotated bibliography. Department of Wildlife Conservation/GEF/UNDP/FAO, Colombo, Sri Lanka. 48 pp.
45. De Silva, A. (2011). Some observations of malformation, eye disease, parasitic and viral infection and the effects of agrochemicals on amphibians in Sri Lanka. FrogLog, 98: 24–25.26.
46. De Silva, M. (2006). An experimental phytotelm Conservation project for a microhylid frog Ramanella nagaoi. Loris, 24: 15–19.
47. De Silva, S. and Wijayathilaka N. (2019). Bioacoustics of Sri Lankan amphibians: a review of current knowledge and conservation significance. Journal of Tropical Forestry and Environment, 9 (1): 1–6. https://doi.org/10.31357/jtfe.v9i1.3944 [DOI]
48. Dinesh, K. P., Radhakrishnan, C., Channakeshavamurthy, B. H. and Kulkarni, N. U. (2015). Checklist of Amphibia of India, updated till January 2015 available at http://mhadeiresearchcenter.org/resources (Accessed 22.12.2016).
49. Dubois, A. (1983). Note préliminaire sur le groupe de Rana (Tomopterna) breviceps Schneider, 1799 (Amphibiens, Anoures), avec diagnose d'une sous-expèce nouvelle de Ceylan. Alytes, 2 (4): 163–170.
50. Dubois, A. and Ohler, A. (2001). A new genus for an aquatic ranid (Amphibia, Anura) from Sri Lanka. Alytes, 19 (2–4): 81–106.
51. Duméril, A. M. C. and Bibron, G. (1841). Erpétologie Genérale ou Histoire Naturelle Complète des Reptiles. Volume 8. Librarie Enclyclopedique de Roret, Paris. 792 pp. https://doi.org/10.5962/bhl.title.45973 [DOI]
52. Dutta, S. K. (1997). Amphibians of India and Sri Lanka (Checklist and Bibliography). Odyssey Publishing House, Orissa. xiii+342+xxii pp.
53. Dutta, S. K. and Manamendra-Arachchi, K. (1996). The Amphibian fauna of Sri Lanka. Wildlife Heritage Trust of Sri Lanka, Colombo, Sri Lanka. 232 pp.
54. Fernando, P. and Siriwardhane, M. (1996). Microhyla karunaratnei (Anura: Microhylidae), a new species of frog endemic to Sri Lanka. Journal of South Asian Natural History, 2 (1): 135–142.
55. Fernando, P., Dayawansa, N. and Siriwardhane, M. (1994). Bufo kotagamai, a new toad (Bufonidae) from Sri Lanka. Journal of South Asian Natural History, 1 (1): 119–124.
56. Fernando, S. S., Wickramasingha, L. J. M. and Rodirigo, R. K. (2007). A new species of endemic frog belonging to genus Nannophrys Günther, 1869 (Anura: Dicroglossinae) from Sri Lanka. Zootaxa, 1403 (1): 55–68. http://dx.doi.org/10.11646/zootaxa.1403.1.3 [DOI]
57. Ferraz, G., Russell, G. J., Stouffer, P. C., Bierregaard, R. O., Pimm, S. L. and Lovejoy, T. E. (2003). Rates of species loss from Amazonian forest fragments. PNAS, 100 (24): 14069–14073. https://doi.org/10.1073/pnas.2336195100 [DOI]
58. Fitzinger, L. J. F. J. (1826). Neue Classification der Reptilien nach ihren Natürlichen Verwandtschaften nebst einer Verwandtschafts-Tafel und einem Verzeichnisse der Reptilien-Sammlung des K. K. Zoologisch Museum’s zu Wien. J. G. Heubner, Wien. 66 pp. https://doi.org/10.5962/bhl.title.4683 [DOI]
59. Fitzinger, L. J. F. J. (1843). Systema Reptilium. Fasciculus Primus. Braumüller ET Seidel, Wien. 106 pp. https://doi.org/10.5962/bhl.title.4694 [DOI]
60. Frost, D. R. (2018). Amphibian Species of the World: an Online Reference. Version 6.0. Electronic Database available at http://research.amnh.org/herpetology/amphibia/index.html. American Museum of Natural History, New York, USA (Accessed 21.09.2018).
61. Frost, D. R., Grant, T., Faivovich, J., Bain, R. H., Haas, A., Haddad, C. F. B., De Sá, R. O., Channing, A., Wilkinson, M., Donnellan, S. C., Raxworthy, C. J., Campbell, J. A., Blotto, B. L., Moler, P., Drewes, R. C., Nussbaum, R. A., Lynch, J. D., Green, D. M. and Wheeler, W. C. (2006). The amphibian tree of life. Bulletin of the American Museum of Natural History, 297: 1–370. Available at http://digitallibrary.amnh.org/dspace/handle/2246/5781 (Accessed 02.12.2018).
62. Garg, S., Senevirathne, G., Wijayathilaka, N., Phuge, S., Deuti, K., Manamendra-Arachchi, K., Meegaskumbura, M. and Biju, S. D. (2018). An integrative taxonomic review of the South Asian microhylid genus Uperodon. Zootaxa, 4384 (1): 1–88. https://doi.org/10.11646/zootaxa.4384.1.1 [DOI]
63. Gosz, J. R. (1993). Ecotone hierarchies. Ecological Applications, 3 (3): 369–376. https://doi.org/10.2307/1941905 [DOI]
64. Gower, D. J., Bahir, M. M., Mapatuna, Y., Pethiyagoda, R., Raheem, D. and Wilkinson, M. (2005). Molecular phylogenetics of Sri Lankan Ichthyophis (Amphibia: Gymnophiona: Ichthyophiidae), with discovery of a cryptic species. The Raffles Bulletin of Zoology, 12: 153–161.
65. Gravenhorst, J. L. C. (1829). Deliciae Musei Zoologici Vratislaviensis. Fasciculus primus. Chelonios ET Batrachia. Leopold Voss, Leipzig. 106 pp.
66. Gray, J. E. (1825). A synopsis of the genera of reptiles and amphibia, with a description of some new species. Annals of Philosophy, New Series, 10: 193–217.
67. Gray, J. E. (1834). Illustrations of Indian Zoology; chiefly selected from the Collection of Major-General Hardwicke, F. R. S. Volume 1. Part 3. Treuttel, Wurtz, Treuttel, Jun. and Richter, London. 100 pls. https://doi.org/10.5962/bhl.title.95127 [DOI]
68. Gunatilleke, I. A. U. N., Gunatilleke, C. V. S. and Dilhan, M. A. A. B. (2005). Plant biogeography and conservation of the south-western hill forests of Sri Lanka. The Raffles Bulletin of Zoology, 12: 9–22.
69. Günther, A. C. L. G. (1858). Neue Batrachier in der Sammlung des britischen Museums. Archiv für Naturgeschichte Berlin, 24: 319–328. https://doi.org/10.5962/bhl.part.5288 [DOI]
70. Günther, A. C. L. G. (1859) [1858]. Catalogue of the Batrachia Salientia in the Collection of the British Museum. London: Taylor and Francis. xvi+160 pp. https://doi.org/10.5962/bhl.title.20903 [DOI]
71. Günther, A. C. L. G. (1864). The reptiles of British India. London, Ray Society. xxvii+452 pp. https://doi.org/10.5962/bhl.title.5012 [DOI]
72. Günther, A. C. L. G. (1869) [1868]. First account of species of tailless batrachians added to the collection of the British Museum. Proceedings of the Zoological Society of London, 1868: 478–490.
73. Günther, A. C. L. G. (1872). Descriptions of some Ceylonese reptiles and batrachians. Annals and Magazine of Natural History, (4) 9: 85–88.
74. Günther, A. C. L. G. (1876a) [1875]. Third report on collections of Indian reptiles obtained by the British Museum. Proceedings of the Zoological Society of London, 1875: 567–577.
75. Günther, A. C. L. G. (1876b). Notes on the mode of propagation of some Ceylonese tree-frogs, with description of two new species. Annals and Magazine of Natural History, (4) 17: 377–380.
76. Gurushankara, H. P., Krishnamurthy, S. V. and Vasudev, V. (2007). Morphological abnormalities in natural populations of common frogs inhabiting agroecosystems of central Western Ghats. Applied Herpetology, 4 (1): 39–45. https://doi.org/10.1163/157075407779766651 [DOI]
77. Hu, J., Li, C., Xie, F. and Jiang, J. (2012). Endemic amphibians and their distribution in China. Asian Herpetological Research, 3 (2): 163–171. https://doi.org/10.3724/SP.J.1245.2012.00163 [DOI]
78. Inger, R. F. (1972). Bufo of Eurasia. In: Blair, W.F. (Ed.), Evolution in the genus Bufo. University of Texas Press, Austin. pp. 102–118.
79. IUCN (International :union: for Conservation of Nature) Sri Lanka (1999). The 1999 Red List of Threatened Fauna and Flora of Sri Lanka. IUCN Sri Lanka, Colombo. 113 pp.
80. IUCN Sri Lanka (2007). The Fauna of Sri Lanka (Status of taxonomy, research and conservation). IUCN Sri Lanka, Colombo. 148 pp.
81. IUCN Sri Lanka (2012). The Fauna of Sri Lanka (Status of taxonomy, research and conservation). IUCN Sri Lanka, Colombo. 452 pp.
82. Janzen, P. and Bopage, M. (2011). The herpetofauna of a small and unprotected patch of tropical rainforest in Morningside, Sri Lanka. Amphibian and Reptile Conservation, 5 (2): 1–13.
83. Jayawardena, B., Senevirathne, G., Wijayathilaka, N., Ukuwela, K., Manamendra-Arachchi, K. and Meegaskumbura, M. (2017). Species boundaries, biogeography and evolutionarily significant units in dwarf toads: Duttaphrynus scaber and D. atukoralei (Bufonidae: Adenominae). Ceylon Journal of Science, 46: 79–87. http://doi.org/10.4038/cjs.v46i5.7455 [DOI]
84. Jerdon, T. C. (1853). Catalogue of reptiles inhabiting the Peninsula of India. The Journal of the Asiatic Society of Bengal, 22: 522–534.
85. Johnson, P. T. J., Chase, J. M., Dosch, K. L., Hartson, R. B., Gross, J. A., Larson, D. J., Sutherland, D. R. and Carpenter, S. R. (2007). Aquatic eutrophication promotes pathogenic infection in amphibians. PNAS, 104 (40): 15781–15786. https://doi.org/10.1073/pnas.0707763104 [DOI]
86. Joshy, S. H., Alam, M. S., Kurabayashi, A., Sumida, M. and Kuramoto, M. (2009). Two new species of the genus Euphlyctis (Anura, Ranidae) from southwestern India, revealed by molecular and morphological comparisons. Alytes, 26 (1–4): 97–116.
87. Kaiser, K. and Hammers, J. L. (2009). The effect of anthropogenic noise on male advertisement call rate in the Neotropical treefrog, Dendropsophus triangulum. Behaviour, 146 (8): 1053–1069.
88. Kaiser, K., Scofield, D. G., Alloush, M., Jones, R. M., Marczak, S., Martineau, K., Oliva, M. A. and Narins, P. M. (2011). When sounds collide: the effect of anthropogenic noise on a breeding assemblage of frogs in Belize, Central America. Behaviour, 148 (2): 215–232. https://doi.org/10.1163/000579510X551660 [DOI]
89. Kandamby, D. S. (2001). Some amphibians observed in Galle District, Sri Lanka. Lyriocephalus, 4: 89–92.
90. Kandamby, D. S. and Batuwita, S. (2001). Some observations on the distribution of Rhacophorus cavirostris (Gunther, 1868) endemic tree frog from Sri Lanka. Lyriocephalus, 4: 93–94.
91. Kark, S. and van Rensburg, B. J. (2006). Ecotones: Marginal or central areas of transition? Journal of Ecology and Evolution, 52 (1): 29–53. https://doi.org/10.1560/IJEE.52.1.29 [DOI]
92. Karunarathna, D. M. S. S., Abeywardena, U. T. I., Asela, M. D. C. and Kekulandala, L. D. C. B. (2008). A preliminary survey of the amphibian fauna in Nilgala Forest area and its vicinity, Monaragala District, Sri Lanka. Herpetological Conservation and Biology, 3 (2): 264–272.
93. Karunarathna, S., Bauer, A. M., De Silva, A., Surasinghe, T., Somaratna, L., Madawala, M., Gabadage, D., Botejue, M., Henkanaththegedara, S. and Ukuwela, K. D. B. (2019a). Description of a new species of the genus Cnemaspis Strauch, 1887 (Reptilia: Squamata: Gekkonidae) from the Nilgala Savannah forest, Uva Province of Sri Lanka. Zootaxa, 4545 (3): 389–407. https://doi.org/10.11646/zootaxa.4545.3.4 [DOI]
94. Karunarathna, S., Poyarkov, N. A., De Silva, A., Madawala, M., Botejue, M., Gorin, V. A., Surasinghe, T., Gabadage, D., Ukuwela, K. D. B. and Bauer, A. M. (2019b). Integrative taxonomy reveals six new species of day geckos of the genus Cnemaspis Strauch, 1887 (Reptilia: Squamata: Gekkonidae) from geographically-isolated hill forests in Sri Lanka. Vertebrate Zoology, 69(3): 247–298. http://doi.org/10.26049/VZ69-3-2019-02 [DOI]
95. Karunarathna, S., Ranwala, S., Surasinghe, T. and Madawala, M. (2017). Impact of vehicular traffic on vertebrate fauna in Horton Plains and Yala national parks of Sri Lanka: some implications for conservation and management. Journal of Threatened Taxa, 9 (3): 9928–9939. http://doi.org/10.11609/jott.2715.93.9928-9939 [DOI]
96. Karunarathna, S., Henkanaththegedara, S., Gabadage, D., Botejue, M., Madawala, M. and Surasinghe, T. D. (2016). Ecology and demography of the critically endangered Kandian torrent toad Adenomus kandianus: a long-lost endemic species of Sri Lanka. Oryx, 51 (4): 619–626. https://doi.org/10.1017/S0030605316000594 [DOI]
97. Kelaart, E. F. (1853). Prodromus faunae zeylanicae; being contributions to the zoology of Ceylon. Published by the author, Colombo, Sri Lanka. vii+xxxiii+197+62+iv pp
98. Kelaart, E. F. (1854). Prodromus faunae zeylanicae; being contributions to the zoology of Ceylon. Government Press, Colombo, Sri Lanka. 23+31+xvi pp. https://doi.org/10.5962/bhl.title.123402 [DOI]
99. Khajeh, A., Mohammadi, Z., Ghorbani, F., Meshkani, J., Rastegar-Pouyani, E. and Torkamanzehi, A. (2014). New insights into the taxonomy of the skittering frog Euphlyctis cyanophlyctis complex (Schneider, 1799) (Amphibia: Dicroglossidae) based on mitochondrial 16S rRNA gene sequences in southern Asia. Acta Herpetologica, 9 (2): 159–166. https://doi.org/10.13128/Acta_Herpetol-14013 [DOI]
100. Kirtisinghe, P. (1946). The genus Nannophrys Günther (Amphibia, Ranidae) with the description of a new species. Ceylon Journal of Science. Section B, Zoology, 23: 105–107.
101. Kirtisinghe, P. (1957). The Amphibia of Ceylon. Published by the author, Colombo, Sri Lanka. xiii+112 pp.
102. Kotagama, S. W., Arudpragasam, K. D. and Kotalawala, I. (1981). A check list of Amphibia of Sri Lanka. National Science Council (National Science Foundation), Colombo, Sri Lanka. 8 pp.
103. Kottawa-Arachchi, J. D. and Wijeratne, M. A. (2017). Climate change impacts on biodiversity and ecosystems in Sri Lanka: a review. Nature Conservation Research, 2 (3): 2–22. https://doi.org/10.24189/ncr.2017.042 [DOI]
104. Kuhl, H. and Van Hasselt, J. C. (1822). Uittreksels uit brieven van de Heeren Kuhl en Van Hasselt, aan de Heeren C. J. Temminck, Th. Van Swinderen en W. De Haan. Algemeene Konst- en LetterBode, 7: 99–104.
105. Laurent, R. F. (1943). Contribution a l’osteologie et a la systematique des rhacophorides non Africains. Bulletin du Musée Royal d'Histoire Naturelle de Belgique, 19: 1–16.
106. Legg, C. and Jewell, N. (1995). A 1:50,000-scale forest map of Sri Lanka: the basis for a national forest geographic information system. Sri Lanka Forester, 20 (3, 4): 35–44.
107. Lesson, R.-P. (1834). Reptiles. In: Bélanger, C. (ed.), Voyage aux Indes-Orientales, par le nord de l'Europe, les provinces du Caucase, la Géorgie, l'Arménie et la Perse, suivi de details topographiques, statistiques et autres sur le Pégou, les Isles de Java, de Maurice et de Bourbon, sur le Cap-de-bonne-Espérance et Sainte-Hélène, pendant ... 1825–29. Zoologie (Texte) [Octavo] and Atlas [Quarto], Paris, Arthus Bertrand, 291–336 pp.
108. Li, J. T., Li, Y., Klaus, S., Rao, D. Q., Hillis, D. M. and Zhang, Y. P. (2013). Diversification of rhacophorid frogs provides evidence for accelerated faunal exchange between India and Eurasia during the Oligocene. PNAS, 110 (9): 3441–3446. https://doi.org/10.1073/pnas.1300881110 [DOI]
109. Lichtenstein, H. and Martens, E. (1856). Nomenclator Reptilium et Amphibiorum Musei Zoologici Berolinensis. Namenverziechniss der in der zoologischen Sammlung der Königlichen Universität zu Berlin aufgestellten Arten von Reptilien und Amphibien nach ihren Ordnungen, Familien und Gattungen. Berlin. iv+48 pp.
110. Linnaeus, C. (1758). Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. 10th Edition. Volume 1. L. Salvii, Stockholm. 775 pp. https://doi.org/10.5962/bhl.title.37256 [DOI]
111. Lütken, C. F. (1864) [1863]. Nogle ny Krybyr og Padder. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, (2)4: 292–311.
112. Maduwage, K. P., Silva, A. and Batuwita, S. (2003). Snakes killed on the Roads of the Gardens of University of Peradeniya. Loris, 23: 22–24.
113. Manamendra-Arachchi, K. and Gabadage, D. E. (1996). Limnonectes kirtisinghei, a new species of ranid frog from Sri Lanka. Journal of South Asian Natural History, 2: 31–42.
114. Manamendra-Arachchi, K. and Pethiyagoda, P. (1998). A synopsis of the Sri Lankan Bufonidae (Amphibia: Anura) with description of new species. Journal of South Asian Natural History, 3: 213–248.
115. Manamendra-Arachchi, K. and Pethiyagoda, R. (2001a). Ramanella nagaoi, a new tree-hole frog (Microhylidae) from southern Sri Lanka. Journal of South Asian Natural History, 5: 121–133.
116. Manamendra-Arachchi, K. and Pethiyagoda, R. (2001b). Polypedates fastigo, a new tree frog (Ranidae: Rhacophorinae) from Sri Lanka. Journal of South Asian Natural History, 5: 191–199.
117. Manamendra-Arachchi, K. and Pethiyagoda, R. (2005). The Sri Lankan shrub-frogs of the genus Philautus Gistel, 1848 (Ranidae: Rhacophorinae), with description of 27 new species. The Raffles Bulletin of Zoology, 12: 163–303.
118. Manamendra-Arachchi, K. and Pethiyagoda, R. (2006). Amphibians of Sri Lanka (in Sinhala). Wildlife Heritage Trust of Sri Lanka, Colombo, Sri Lanka. 440 pp.
119. Meegaskumbura, M. and Manamendra-Arachchi, K. (2005). Descriptions of eight new species of shrub frogs (Ranidae: Rhacophorinae: Philautus) from Sri Lanka. The Raffles Bulletin of Zoology, 12: 305–338.
120. Meegaskumbura, M. and Manamendra-Arachchi, K. (2011). Two new species of shrub frogs (Rhacophoridae: Pseudophilautus) from Sri Lanka. Zootaxa, 2747 (1): 1–18. http://dx.doi.org/10.11646/zootaxa.2747.1.1 [DOI]
121. Meegaskumbura, M., Bossuyt, F., Pethiyagoda, R., Manamendra-Arachchi, K., Bahir, M. M., Milinkovitch, M. C. and Schneider, C. J. (2002). Sri Lanka: an amphibian hotspot. Science, 298: 379. https://doi.org/10.1126/science.298.5592.379 [DOI]
122. Meegaskumbura, M., Manamendra-Arachchi, K. and Pethiyagoda, R. (2009). Two new species of shrub frogs (Rhacophoridae: Philautus) from the lowlands of Sri Lanka. Zootaxa, 2122 (1): 51–68. http://dx.doi.org/10.11646/zootaxa.2747.1.1 [DOI]
123. Meegaskumbura, M., Manamendra-Arachchi, K., Bowatte, G. and Meegaskumbura, S. (2012a). Rediscovery of Pseudophilautus semiruber, a diminutive shrub frog (Rhacophoridae: Pseudophilautus) from Sri Lanka. Zootaxa, 3229 (1): 58–68. http://dx.doi.org/10.11646/zootaxa.3229.1.4 [DOI]
124. Meegaskumbura, M., Meegaskumbura, S., Gunatilleke, N., Manamendra-Arachchi, K., Bowatte, G., Gunatilleke, T. and Bandara, C. (2012b). Conservation and biogeography of threatened Amphibians of Eastern Sinharaja. FrogLog, 100: 12–14.
125. Meegaskumbura, M., Meegaskumbura, S., Bowatte, G., Manamendra-Arachchi, K., Pethiyagoda, R., Hanken, J. and Schneider, C. J. (2010). Taruga (Anura: Rhacophoridae), a new genus of foam-nesting tree frogs endemic to Sri Lanka. Ceylon Journal of Science (Biological Sciences), 39 (2): 75–94. http://doi.org/10.4038/cjsbs.v39i2.2995 [DOI]
126. Meegaskumbura, M., Manamendra-Arachchi, K., Schneider, C. J. and Pethiyagoda, R. (2007). New species amongst Sri Lanka’s extinct shrub frogs (Amphibia: Rhacophoridae: Philautus). Zootaxa, 1397 (1): 1–15. http://dx.doi.org/10.11646/zootaxa.1397.1.1 [DOI]
127. Meegaskumbura, M., Seneviratne, G., Wijayathilaka, N., Jayawardena, B., Bandara, C., Manamendra-Arachchi, K. and Pethiyagoda, R. (2015). The Sri Lankan torrent toads (Bufonidae: Adenominae: Adenomus): species boundaries assessed using multiple criteria. Zootaxa, 3911 (2): 245–261. http://dx.doi.org/10.11646/zootaxa.3911.2.6 [DOI]
128. Meegaskumbura, M., Seneviratne, G., Manamendra-Arachchi, K., Pethiyagoda, R., Hanken, J. and Schneider, C. J. (2019). Diversification of shrub frogs (Rhacophoridae, Pseudophilautus) in Sri Lanka-Timing and geographic context. Molecular Phylogenetics and Evolution, 132: 14–24. https://doi.org/10.1016/j.ympev.2018.11.004 [DOI]
129. Méhely, L. (1897). Zur Herpetologie von Ceylon. Természetrajzi Füzetek, 20: 55–70.
130. Min, M. S., Lee, H. and Waldman, B. (2011). Amphibian Population Declines and Chytridiomycosis in South Korea. FrogLog, 98: 12–13.
131. Mittermeier, R. A., Myers, N., Thomsen, J. B., Da Fonseca, G. A. B. and Olivieri, S. (1998). Biodiversity hotspots and major tropical wilderness areas: approaches to setting conservation priorities. Conservation Biology, 12 (3): 516–520. https://doi.org/10.1046/j.1523-1739.1998.012003516.x [DOI]
132. Müller, F. (1887). Fünfter Nachtrag zum Katalog der herpetologischen Sammlung des Basler Museum. Verhandlungen der Naturforschenden Gesellschaft in Basel, 8: 249–296.
133. Myers, N., Mittermeier, R. A., Mittermeier, C. G., da Fonseca, G. A. B. and Kent, J. (2000). Biodiversity hotspots for conservation priorities. Nature, 403: 853–858. https://doi.org/10.1038/35002501 [DOI]
134. Naniwadekar, R. and Vasudevan, K. (2007). Patterns in diversity of anurans along an elevational gradient in the Western Ghats, South India. Journal of Biogeography, 34 (5): 842–853. https://doi.org/10.1111/j.1365-2699.2006.01648.x [DOI]
135. Nguyen, T. T., Nguyen, T. V., Ziegler, T., Pasmans, F. and Martel, A. (2017). Trade in wild anurans vectors the urodelan pathogen Batrachochytrium salamandrivorans into Europe. Amphibia-Reptilia, 38 (4): 554–556. https://doi.org/10.1163/15685381-00003125 [DOI]
136. Nishikawa, K., Matsui, M., Yong, H.-S., Ahmad, N., Yambun, P., Belabut, D. M., Sudin, A., Hamidy, A., Orlov, N. L., Ota, H., Yoshikawa, N., Tominaga, A. and Shimada, T. (2012). Molecular phylogeny and biogeography of caecilians from Southeast Asia (Amphibia, Gymnophiona, Ichthyophiidae), with special reference to high cryptic species diversity in Sundaland. Molecular Phylogenetics and Evolution, 63 (3): 714–723. https://doi.org/10.1016/j.ympev.2012.02.017 [DOI]
137. Nussbaum, R. A. and Gans, C. (1980). On the Ichthyophis (Amphibia: Gymnophiona) of Sri Lanka. Spolia Zaylanica, 35 (1, 2): 137–154.
138. Odum, E. P. (1953). Fundamentals of Ecology. First Edition. W. B. Saunders Company, Philadelphia. Xii + 384 pp.
139. Parker, H. W. (1934). A monograph of the frogs of the family Microhylidae. The British Museum (Natural History), London. 208 pp.
140. Parker, H. W. and Osman Hill, W. C. (1948). Frogs of the genus Microhyla from Ceylon. Annals and Magazine of Natural History, 1 (10): 759–764. https://doi.org/10.1080/00222934808653944 [DOI]
141. Peabotuwage, I., Bandara, I. N., Samarasinghe, D., Perera, N., Madawala, M., Amarasinghe, C., Kandambi, H. K. D. and Karunarathna, D. M. S. S. (2012). Range extension for Duttaphrynus kotagamai (Amphibia: Bufonidae) and a preliminary checklist of herpetofauna from the Uda Maliboda Trail in Samanala Nature Reserve, Sri Lanka. Amphibian and Reptile Conservation, 5 (2): 52–64.
142. Peloso, P. L. V., Frost, D. R., Richards, S. J., Rodrigues, M. T., Donnellan, S., Matsui, M., Raxworthy, C. J., Biju, S. D., Lemmon, E. M., Lemmon, A. R. and Wheeler, W. C. (2016). The impact of anchored phylogenomics and taxon sampling on phylogenetic inference in narrow-mouthed frogs (Anura, Microhylidae). Cladistics, 32 (2): 113–140. https://doi.org/10.1111/cla.12118 [DOI]
143. Perl, R. G. B., Geffen, E., Malka, Y., Barocas, A., Renan, S., Vences, M. and Gafny, S. (2018). Population genetic analysis of the recently rediscovered Hula painted frog (Latonia nigriventer) reveals high genetic diversity and low inbreeding. Scientific Reports, 8: 5588. https://doi.org/10.1038/s41598-018-23587-w [DOI]
144. Peters, W. C. H. (1863). Über eine neue Schlangen-gattung, Styporhynchus, und verschiedene andere Amphibien des zoologischen Museum. Monatsberichte der Königlichen Preussische Akademie des Wissenschaften zu Berlin, 1863: 399–413.
145. Pethiyagoda, R. (1991). Freshwater fishes of Sri Lanka. Wildlife Heritage Trust of Sri Lanka, Colombo, Sri Lanka. 362 pp.
146. Pethiyagoda, R. and Manamendra-Arachchi, K. (1998). Evaluating Sri Lanka’s amphibian diversity. Occassional Papers of the Wildlife Heritage Trust, 2: 1–12.
147. Pounds, J. A. and Puschendorf, R. (2004). Clouded futures. Nature, 427: 107–109. https://doi.org/10.1038/427107a [DOI]
148. Rajapakshe, R. V. P. J. and De Silva, A. (2001). Preliminary observations of the parasites of amphibian fauna at Horton Plains National Park. Lyriocephalus, 4: 166–167.
149. Risser, P. G. (1995). The status of the science examining ecotones: a dynamic aspect of landscape is the area of steep gradients between more homogeneous vegetation associations. BioScience, 45 (5): 318–325. https://doi.org/10.2307/1312492 [DOI]
150. Ryan, M. J., Scott, N. J., Cook, J. A., Willink, B., Chaves, G., Bolãnos, F., García-Rodríguez, A., Latella, I. M. and Koerner, S. E. (2015). Too wet for frogs: changes in a tropical leaf litter community coincide with La Niña. Ecosphere, 6 (1): 1–10. http://dx.doi.org/10.1890/ES14-00352.1 [DOI]
151. Sanchez, E., Biju, S. D., Islam, M. M., Hasan, M., Ohler, A., Vences, M. and Kurabayashi, A. (2018). Phylogeny and classification of fejervaryan frogs (Anura: Dicroglossidae). Salamandra, 54 (2): 109–116.
152. Schilthuizen, M. (2000). Ecotone: Speciation-prone. Trends in Ecology and Evolution, 15 (4): 130–131. https://doi.org/10.1016/S0169-5347(00)01839-5 [DOI]
153. Schneider, J. G. (1799). Historia Amphibiorum Naturalis et Literarariae. Fasciculus Primus. Continens Ranas, Calamitas, Bufones, Salamandras et Hydros in Genera et Species Descriptos Notisque suis Distinctos. Friederici Frommanni, Jena. 302 pp.
154. Seale, D. B. (1980). Influence of amphibian larvae on primary production, nutrient flux, and competition in a pond ecosystem. Ecology, 61 (6): 1531–1550. https://doi.org/10.2307/1939059 [DOI]
155. Seba, A. (1735). Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam: Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, et depingendum curavit. Volume II. Wetstenium, Smith and Janssonio–Waesbergios, Amsterdam. 1–32+1–154 pp., 114 pls. https://doi.org/10.5962/bhl.title.62760 [DOI]
156. Senanayake, F. R., Soulè, M. and Senner, J. W. (1977). Habitat values and endemicity in the vanishing rains forests of Sri Lanka. Nature, 265: 351–354. https://doi.org/10.1038/265351a0 [DOI]
157. Senevirathne, G., Samarawickrama, V. A. M. P. K., Wijayathilaka, N., Manamendra-Arachchi, K., Bowatte, G., Samarawickrama, D. R. N. S. and Meegaskumbura, M. (2018). A new frog species from rapidly dwindling cloud forest streams of Sri Lanka-Lankanectes pera (Anura, Nyctibatrachidae). Zootaxa, 4461 (4): 519–538. http://dx.doi.org/10.11646/zootaxa.4461.4.4 [DOI]
158. Shaw, G. (1802). General Zoology or Systematic Natural History. Volume III, Part 1. Amphibia. Thomas Davison, London. 312 pp.
159. Seymour, R. S. (1999). Respiration of aquatic and terrestrial amphibian embryos. American Zoologist, 39 (2): 261–270. www.jstor.org/stable/3884249
160. Shreve, B. (1940). A new Rhacophorus and a new Philautus from Ceylon. Proceedings of the Biological Society of Washington, 53: 105–108.
161. Silva, P. and Kurukulasuriya, M. (2010). Invasive alien fauna in Sri Lanka - Introduction, spread, impacts and management, In: Marambe, B., Silva, P., Wijesundara, S. and Atapattu, N. (Eds.), Invasive Alien Species in Sri Lanka- strengthening capacity to control their introduction and spread. Biodiversity Secretariat of the Ministry of Environment, Colombo, Sri Lanka. pp. 39–61.
162. Sredl, M. J. and Collins, J. P. (1992). The interaction of predation, competition, and habitat complexity in structuring an amphibian community. Copeia, 1992 (3): 607–614.
163. Stuart, S. N., Chanson, J. S., Cox, N. A., Young, B. E., Rodrigues, A. S. L., Fischman, D. L. and Waller, R.W. (2004). Status and trends of amphibian declines and extinctions worldwide. Science, 306 (5702): 1783–1786. http://dx.doi.org/10.1126/science.1103538 [DOI]
164. Sun, J. W. C. and Narins, P. M. (2005). Anthropogenic sounds differentially affect amphibian call rate. Biological Conservation, 121 (3): 419–427. https://doi.org/10.1016/j.biocon.2004.05.017 [DOI]
165. Surasinghe, T. D. (2009). Conservation and distribution status of amphibian fauna in Sri Lanka. Biodiversity, 10 (1): 3–17. https://doi.org/10.1080/14888386.2009.9712632 [DOI]
166. Taigen, T. L., Pough, F. H. and Stewart, M. M. (1984). Water balance of terrestrial anuran (Eleutherodactylus coqui) eggs: importance of parental care. Ecology, 65 (1): 248–255. https://doi.org/10.2307/1939477 [DOI]
167. Taylor, B., Skelly, D., Demarchis, L. K., Slade, M. D., Galusha, D. and Rabinowitz, P. M. (2005). Proximity to pollution sources and risk of amphibian limb malformation. Environmental Health Perspectives, 113 (11): 1497–1501. https://doi.org/10.1289/ehp.7585 [DOI]
168. Taylor, E. H. (1965). New Asiatic and African caecilians with redescriptions of certain other species. The University of Kansas Science Bulletin, 46 (6): 253–302. https://doi.org/10.5962/bhl.part.20077 [DOI]
169. Taylor, E. H. (1969). Miscellaneous notes and descriptions of new forms of caecilians. The University of Kansas Science Bulletin, 48 (9): 281–296. https://doi.org/10.5962/bhl.part.11223 [DOI]
170. Tschudi, J. J. (1838). Classification der Batrachier, mit Berucksichtigung der fossilen Thiere dieser Abtheilung der Reptilien. Neuchatel, Petitpierre. ii+99 pp., 6 pls. https://doi.org/10.5962/bhl.title.4883 [DOI]
171. UNESCO (United Nations Educational, Scientific and Cultural Organization) (2010). UNESCO Headquarters, 7, Place de Fontenoy, 75352, Paris, 07 SP, France. http://whc.unesco.org/en/list/1203/documents (Accessed 17 November 2019).
172. Van Bocxlaer, I., Biju, S. D., Loader, S. P. and Bossuyt, F. (2009). Toad radiation reveals into-India dispersal as a source of endemism in the Western Ghats-Sri Lanka biodiversity hotspot. BMC Evolutionary Biology, 9, 131: 1–10. https://doi.org/10.1186/1471-2148-9-131 [DOI]
173. WCSG (Wildlife Conservation Society-Galle) (2008). The study of the faunal diversity in Galle District- Southern, Sri Lanka. Galle, Wildlife Conservation Society- Galle, Sri Lanka. 44 pp.
174. WCSG (Wildlife Conservation Society-Galle) (2009). The study of the faunal diversity in Matara District- Southern, Sri Lanka. Wildlife Conservation Society- Galle, Sri Lanka. 43 pp.
175. Weerawardhena, S. R. and Russell, A. (2012). Cover-dependency of anurans in the Riverstone, Knuckles mountain forest range, Sri Lanka. Taprobanica, 4 (1): 12–19.
176. Werner, W. L. (1988). Canopy dieback in the upper montane rain forests of Sri Lanka. GeoJournal, 17: 245–248. https://doi.org/10.1007/BF02432929 [DOI]
177. Wickramasinghe, L. J. M. and Munindradasa, D. A. I. (2007). Review of the genus Cnemaspis Strauch, 1887 (Sauria: Gekkonidae) in Sri Lanka with the description of five new species. Zootaxa, 1490 (1): 1–63. http://dx.doi.org/10.11646/zootaxa.1490.1.1 [DOI]
178. Wickramasinghe, L. J. M., Bandara, I. N., Vidanapathirana, D. R., Tennakoon, K. H., Samarakoon, S. R. and Wickramasinghe, N. (2015). Pseudophilautus dilmah, a new species of shrub frog (Amphibia: Anura: Rhacophoridae) from a threatened habitat Loolkandura in Sri Lanka. Journal of Threatened Taxa, 7: 7089–7110. https://doi.org/10.11609/JoTT.o3501.7089-110 [DOI]
179. Wickramasinghe, L. J. M., Munindradasa, D. A. I. and Fernando, P. (2012a). A new species of Polypedates Tschudi (Amphibia, Anura, Rhacophoridae) from Sri Lanka. Zootaxa, 3498 (1): 63–80. http://dx.doi.org/10.11646/zootaxa.4691.5.6 [DOI]
180. Wickramasinghe, L. J. M., Vidanapathirana, D. R. and Wickramasinghe, N. (2012b). Back from the dead: the world’s rarest toad Adenomus kandianus rediscovered in Sri Lanka. Zootaxa, 3347 (1): 63–68. http://dx.doi.org/10.11646/zootaxa.3347.1.3 [DOI]
181. Wickramasinghe, L. J. M., Vidanapathirana, D. R., Ariyarathne, S., Rajeev, G., Chanaka, A., Pastorini, J., Chathuranga, G. and Wickramasinghe, N. (2013b). Lost and found: One of the world’s most elusive amphibian Pseudophilautus stellatus (Kelaart, 1853) rediscovered. Zootaxa, 3620 (1): 112–128. http://dx.doi.org/10.11646/zootaxa.3620.1.5 [DOI]
182. Wickramasinghe, L. J. M., Vidanapathirana, D. R., Rajeev, G. and Wickramasinghe, N. (2013c). Rediscovery of Pseudophilautus hypomelas (Günther, 1876) (Amphibia: Anura: Rhacophoridae) from the Peak Wilderness, Sri Lanka, a species thought to be extinct! Journal of Threatened Taxa, 5: 5181–5193. http://dx.doi.org/10.11609/JoTT.o3547.5181-93 [DOI]
183. Wickramasinghe, L. J. M., Vidanapathirana, D. R., Rajeev, M. D. G., Ariyarathne, S. C., Chanaka, A. W. A., Priyantha, L. L. D., Bandara, I. N. and Wickramasinghe, N. (2013a). Eight new species of Pseudophilautus (Amphibia, Anura, Rhacophoridae) from Sripada World Heritage Site (Peak Wilderness), a local amphibian hotspot in Sri Lanka. Journal of Threatened Taxa, 5: 3789–3920. https://doi.org/10.11609/JoTT.o3099.3789-920 [DOI]
184. Wijayathilaka, N., Garg, S., Senevirathne, G., Karunarathna, N. and Biju, S. D., Meegaskumbura, M. (2016). A new species of Microhyla (Anura: Microhylidae) from Sri Lanka: an integrative taxonomic approach. Zootaxa, 4066 (3): 331–342. http://dx.doi.org/10.11646/zootaxa.4420.2.1 [DOI]
185. Xie, F., Lau, M. W. N., Stuart, S. N., Chanson, J. S., Cox, N. A. and Fischman, D. L. (2007). Conservation needs of amphibians in China: A review. Science in China Series C, 50: 265. http://doi.org/10.1007/s11427-007-0021-5 [DOI]

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